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Definition
Community characteristics – diversity, stratification,
dominance, structure, periodicity, fluctuations,
stability, interdependence, interactions and
succession.
Biotic community – salient features, types.
Concept of biocoenosis.
Community concept, types and hypotheses.
Community succession.
Important species(Foundation, keystone, invasive)
and concepts (ecotone, edge effect, Niche).
summary
Definition -1
• Communities are assemblages of large number of species
that occur together in space and time and all interact
directly or indirectly with each other.

Definition -2
• A community is defined as groups of different organisms
living together in a particular space

Definition -3
• A biotic community comprises plants and animals present in a
particular area.
• E.g. microbial community, plant community, animal
community








plants and animals belonging to different
taxonomic groups are present in a
community.
The abundance of populations and
number of species varies greatly in a
community.
Species richness = no. of species in a
community.
Species evenness = the distribution of
individuals among the species.
Species diversity is the product of richness
and evenness.
All communities have a physical
structure and biological pattern.
 Stratification is a layering or series of
separations that differently distributes the
organisms occurring in an ecosystem.
 Stratifications can be described in terms
of separations between organisms in
space (vertical or horizontal) or
 In terms of separations between
organisms in time (periodicity).

An ecosystem can be stratified in space
either vertically (layers) or horizontally
(concentric circles).
 Stratification of terrestrial communities
reflects the life forms of plants.
 Stratification results in economy of space
and per unit area of the land surface.
 Organisms in a community display daily,
lunar, seasonal or irregular patterns that
separate them in time.

Tropical
rain forest

Coniferous
forest

Deciduous
forest

Thorn
forest

Thorn
scrub

Tall-grass Short-grass
prairie
prairie

Desert
scrub
All the organisms in a community are not
equally important in determining the nature
of that community.
Only a few species generally exert the
major controlling influence on the
community due to their size, number or
activities.
These groups largely control the energy
flow, which are known as ecological
dominants.
In pasture grass and cattle are ecological
dominants.
A community is a composite biotic unit.
The individual populations of different
species depend on each other and on
the abiotic environment for their food,
shelter and reproduction.
A community consists of mainly 3 groups.
Producers= autotrophic organisms
Consumers= herbivores and carnivores
Decomposers= microbes


The rhythmic changes occur in the
activities of organisms.
1. Seasonal periodicity – temperature, rainfall

and photoperiod determine the
reproductive cycle of its inhabitants.
2. Dial periodicity – animals of terrestrial
community active during the day and
inactive during night.
Even within permanent and stable
community, fluctuations in species
abundance and numbers occur due to
the interplay of biotic potential and
environmental resistance.
 It may be regular or cyclic.
 E.g. host-parasite or predator- prey









Community stability is the degree of
fluctuations in the size of the populations
comprising the community.
Environmental stability is the fluctuations in the
abiotic factors of the ecosystem.
Both the types of stability are closely related.
The structure of community has a major role in
determining the degree of environmental
variability.
For e.g. the organisms of a forest floor is
protected from winds, evaporation and
temperature variations by the thick forest
canopy.
A community is a self-sustained unit.
 The interdependence among different
species of a community is found for
food, reproduction and protection.
 Most communities are interdependent,
or share a mutual responsibility for group
survival.



Competition occurs when organisms of the same
or different species attempt to use an ecological
resource(water, nutrients, light, food or space) in
the same place at same time.



Predator –prey interactions



Symbiosis-

different species rely on each other :3 kinds.

– Mutualism- both partners benefit.
– Commensalism- one partner benefits and the other is
unaffected.

– Parasitism- One benefits the other is harmed.


Trophic structures are the feeding relationships
within communities i.e. who's eating whom



The first trophic level is made up of the primary
producers, the organisms that obtain the
energy from inorganic sources.



Primary producers are photosynthetic
organisms; more generally, primary producers
are autotrophs (i.e., they fix CO2)



Consumers are the heterotrophs, i.e., organisms
that obtain their carbon from other organisms.
It is the development of a community.
 It is a directional change in the community structure as a function of
time.
 There two fundamental types of succession: primary and secondary.
Primary succession is the series of community changes which occur
on an entirely new habitat.
 Secondary succession is the series of community changes which
take place on a previously colonized, but disturbed or damaged
habitat.



There is a balanced assemblage of
autotrophic plants and heterotrophic animals
in the trophic structure of a community.
A community is an aggregate of
organisms, which form a distinct ecological
unit.
2. The size of community unit may be large
(forest community) or small (community of
invertebrates).
3. Different community occurs in different
habitats.
4. The composition and character of a
community is an indicator of the type of
environment that is present.
1.
5.The structure of a community is dependent on:
1) the number of species.2) the relative
number individuals in each species.
6.Species diversity indices give quantitative
expression of community structure.
7.Communities are constantly changing. The
replacement of one community by another is
called community succession. The final stage
in community succession is a climax
community.
On the whole a community is considered as a
highly integrated self-contained organic unit.
Minor
communities

Major
communities

• Are those communities more or less
dependent on neighboring
communities for energy.
• They are secondary aggregations within
a major community.

• Are those communities of sufficient size
and completeness of organization.
• They are relatively independent of
resources from adjacent communities.
• They receive only solar energy from the
sun.
A biocoenosis is a biotic community,
coined by Karl Mobius in 1877.
 This concept defines community as the
interacting organisms living together in a
habitat(biotope).


 Zoocoenosis refers to faunal community
 Phytocoenosis refers to floral community
 Microbiocoenosis refers to microbial
community.
Community concept is one of the most
important principles in Ecology.
 This concept defines community as
diverse organisms usually live together in
an orderly manner.
 Victor E. Shelford has defined “the
community as an assemblage of
populations living in a prescribed area or
physical habitat with a definite trophic
organization and metabolic pattern”.







The individualistic hypothesis
was proposed by H.A.
Gleason( American
Ecologist),1917.
This hypothesis depicted a
community as a chance
assemblage of species found
in an area because they have
similar abiotic requirements.
In other words ―vegetation is
continuously variable in
response to a continuously
varying environment‖.
The interactive hypothesis was proposed
by F.E. Clements in 1916.
 According to him, each community is an
assemblage of closely linked species
having mandatory biotic interactions
that cause the community to function as
an integrated unit.

Communities are stable, integrated, and
orderly entities.
 a. Biologically they have a highlypredictable composition.
 b. The diversity and abundance of a
species in a particular community will be
the same before and after a disturbance.
 c. Communities develop by passing
through a series of predictable stages,
culminating in a stable climax community.

Communities are neither stable nor
predictable.
 a. Plant and animal communities are
ephemeral associations of species that
just happen to share similar climatic
requirements.
 b. It is largely a matter of chance
whether a similar community develops in
the same area after a disturbance
occurs.

The number of species
 The relative
abundance of those
species
 The types of species
 The number, types and
strength of interactions
among species (i.e.,
the structure of food
webs)

 1.Tropical









rain forests

They found at equator.
Temperatures and rainfall are high and
variation is low.
Plants grow all year long.
High productivity, and high amounts of
aboveground biomass.
Highest amounts of BIODIVERSITY on Earth.
Extraordinary structural diversity.
A multilayered tree canopy is intermingled
with vines, epiphytes, shrubs and herbs.
 2.Temperate







Forests

Precipitation is high and relatively constant
throughout the year.
This abundance of moisture allows trees to
dominate the landscape.
Plants experience a seasonal period of
dormancy.
Productivity higher than deserts or
grasslands, lower than tropical forests.
Biodiversity is moderate.
 3.

Grasslands

These areas are also called prairies or steppes.
 Precipitation conditions are quite dry.
 Temperatures are moderate but highly SEASONAL.
 Plant growth occurs in wet and warm months
 Grasslands can develop in forested regions if
recurring fires burn out
 Productivity is lower than in forests.
 Grassland soil is highly fertile.

 4.Deserts

Low annual precipitation
 Low productivity
 Individual plants are widely spaced due
to the intense competition for water.
 Desert species must cope with extreme
temperatures and aridity.





Pond community –
there may be
populations of aquatic
plants, herbivorous
organisms, carnivores
and bacteria at the
bottom.
They are
interdependent for
food supplies and form
a specific group.









Species diversity is a key feature of
biological communities.
Species diversity affects the stability and
productivity of communities.
A latitudinal gradient of species diversity
exists for many taxa.
Species diversity declines as latitude
increases.
High diversity in the tropics leads to high
productivity.
Diversity is positively correlated with stability.
Each community type experiences a
characteristic type, frequency, and
severity of disturbance. This is known as a
community’s disturbance regime.
 Disturbance patterns profoundly affect
the composition of communities.
 Examples—Fires in boreal forests and
tree falls in temperate and tropical
forests.

Definition
•Community succession is
defined as a gradual,
continuous and
unidirectional change in
the species composition of
a natural community over
a long time.
All ecosystems change over time
 There are two types of ecosystem changes



Nondirectional directional changes-ReplacementFluctuations



Directional changes-Biogeographic change- Succession



All these types of changes may occur
simultaneously within a single ecosystem
Primary
Community
succession

succession
Secondary
succession

It takes place
in barren
areas

It takes place
in disturbed
areas
Early stages

Middle stages

Final stages

• Low biomass
• Low nutrient levels
• Low Species
diversity
• Short food chain
• High productivity

• Rich biomass
• Rich organic
nutrients
• More species
diversity

• Stable high biomass
• High organic
nutrients & low
productivity
• Complex food
chain
• High species
diversity
&competition
Pioneer
Community/

Sere /
intervening

First community

communities

Climax
community
Forest Succession

Pioneer
Comm.

Climax
Comm.
Increase in
Species composition
And species diversity
Increase in species density
And heterogeneity
Increase in
Organic nutrients
Metabolic stability in
Community metabolism
Foundation species are considered the
―base‖ of a community, having the
greatest influence on its overall structure.
 They are usually the primary producers:
 Foundation species may physically
modify the environment to produce and
maintain habitats that benefit the other
organisms that use them.
 E.g. the photosynthetic corals of the
coral reef.








A species whose presence is key to
maintaining biodiversity within an ecosystem
and to upholding an ecological community’s
structure.
The species may be a strong interactor or a
weak interactor.
Strong interactors are fewer in number and
critical to the ecosystem function.
Elimination of a keystone species may result in
the loss of many other species in a community.
E.g. beavers of Canada, Bengal tiger of India,
African elephants.
Invasive species are non-indigenous
species whose introduction can cause
harm to the community and the
environment.
 Invasive species are often better
competitors than native species,
resulting in population explosions.
 These new species usually overtake the
native populations, driving them to
localized extinctions.

A niche is the role of an organism in a
community (Charles Elton 1927)- functional
niche = functional status of an organism in its
community.
 Joseph Grinnell (1917)referred niche as an
organism’s physical environment- spatial
niche
 Niche is a property of a biotic community.
 Niches includes all the physical, chemical
and biological conditions.
 The niche concept was formulated to study the
role of organisms in community.

An edge is where two or more different
vegetational communities meet. e.g. forestgrass land.
 An ecotone is where two or more
communities not only meet but also
intergrade.
 Edges result from abrupt changes in soil
type, topographic differences, geomorphic
differences and microclimatic changes.
 Ecotones arise from the blending of two or
more vegetational types.

A community is a complex group of
individuals interacting and sharing an
environment.
 Communities can be characterized by
their structure (the types and numbers of
species present) and dynamics (how
communities change over time).








Dr. B. Victor is a highly experienced postgraduate
professor, retired from the reputed educational
institution - St. Xavier’ s College(Autonomous),
Palayamkottai, India-627001.
He was the dean of sciences, assistant controller of
examinations and coordinator several academic
research workshops.
He has more than 32 years of teaching and research
experience
He has taught a diversity of courses and published
45 research articles in reputed national and
international journals.
Send your comments to : bonfiliusvictor@gmail.com
Biological communities

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Biological communities

  • 1.
  • 2. Definition Community characteristics – diversity, stratification, dominance, structure, periodicity, fluctuations, stability, interdependence, interactions and succession. Biotic community – salient features, types. Concept of biocoenosis. Community concept, types and hypotheses. Community succession. Important species(Foundation, keystone, invasive) and concepts (ecotone, edge effect, Niche). summary
  • 3. Definition -1 • Communities are assemblages of large number of species that occur together in space and time and all interact directly or indirectly with each other. Definition -2 • A community is defined as groups of different organisms living together in a particular space Definition -3 • A biotic community comprises plants and animals present in a particular area. • E.g. microbial community, plant community, animal community
  • 4.
  • 5.      plants and animals belonging to different taxonomic groups are present in a community. The abundance of populations and number of species varies greatly in a community. Species richness = no. of species in a community. Species evenness = the distribution of individuals among the species. Species diversity is the product of richness and evenness.
  • 6. All communities have a physical structure and biological pattern.  Stratification is a layering or series of separations that differently distributes the organisms occurring in an ecosystem.  Stratifications can be described in terms of separations between organisms in space (vertical or horizontal) or  In terms of separations between organisms in time (periodicity). 
  • 7. An ecosystem can be stratified in space either vertically (layers) or horizontally (concentric circles).  Stratification of terrestrial communities reflects the life forms of plants.  Stratification results in economy of space and per unit area of the land surface.  Organisms in a community display daily, lunar, seasonal or irregular patterns that separate them in time. 
  • 9. All the organisms in a community are not equally important in determining the nature of that community. Only a few species generally exert the major controlling influence on the community due to their size, number or activities. These groups largely control the energy flow, which are known as ecological dominants. In pasture grass and cattle are ecological dominants.
  • 10. A community is a composite biotic unit. The individual populations of different species depend on each other and on the abiotic environment for their food, shelter and reproduction. A community consists of mainly 3 groups. Producers= autotrophic organisms Consumers= herbivores and carnivores Decomposers= microbes
  • 11.  The rhythmic changes occur in the activities of organisms. 1. Seasonal periodicity – temperature, rainfall and photoperiod determine the reproductive cycle of its inhabitants. 2. Dial periodicity – animals of terrestrial community active during the day and inactive during night.
  • 12. Even within permanent and stable community, fluctuations in species abundance and numbers occur due to the interplay of biotic potential and environmental resistance.  It may be regular or cyclic.  E.g. host-parasite or predator- prey 
  • 13.      Community stability is the degree of fluctuations in the size of the populations comprising the community. Environmental stability is the fluctuations in the abiotic factors of the ecosystem. Both the types of stability are closely related. The structure of community has a major role in determining the degree of environmental variability. For e.g. the organisms of a forest floor is protected from winds, evaporation and temperature variations by the thick forest canopy.
  • 14. A community is a self-sustained unit.  The interdependence among different species of a community is found for food, reproduction and protection.  Most communities are interdependent, or share a mutual responsibility for group survival. 
  • 15.  Competition occurs when organisms of the same or different species attempt to use an ecological resource(water, nutrients, light, food or space) in the same place at same time.  Predator –prey interactions  Symbiosis- different species rely on each other :3 kinds. – Mutualism- both partners benefit. – Commensalism- one partner benefits and the other is unaffected. – Parasitism- One benefits the other is harmed.
  • 16.  Trophic structures are the feeding relationships within communities i.e. who's eating whom  The first trophic level is made up of the primary producers, the organisms that obtain the energy from inorganic sources.  Primary producers are photosynthetic organisms; more generally, primary producers are autotrophs (i.e., they fix CO2)  Consumers are the heterotrophs, i.e., organisms that obtain their carbon from other organisms.
  • 17. It is the development of a community.  It is a directional change in the community structure as a function of time.  There two fundamental types of succession: primary and secondary. Primary succession is the series of community changes which occur on an entirely new habitat.  Secondary succession is the series of community changes which take place on a previously colonized, but disturbed or damaged habitat. 
  • 18.  There is a balanced assemblage of autotrophic plants and heterotrophic animals in the trophic structure of a community.
  • 19. A community is an aggregate of organisms, which form a distinct ecological unit. 2. The size of community unit may be large (forest community) or small (community of invertebrates). 3. Different community occurs in different habitats. 4. The composition and character of a community is an indicator of the type of environment that is present. 1.
  • 20. 5.The structure of a community is dependent on: 1) the number of species.2) the relative number individuals in each species. 6.Species diversity indices give quantitative expression of community structure. 7.Communities are constantly changing. The replacement of one community by another is called community succession. The final stage in community succession is a climax community. On the whole a community is considered as a highly integrated self-contained organic unit.
  • 21. Minor communities Major communities • Are those communities more or less dependent on neighboring communities for energy. • They are secondary aggregations within a major community. • Are those communities of sufficient size and completeness of organization. • They are relatively independent of resources from adjacent communities. • They receive only solar energy from the sun.
  • 22. A biocoenosis is a biotic community, coined by Karl Mobius in 1877.  This concept defines community as the interacting organisms living together in a habitat(biotope).   Zoocoenosis refers to faunal community  Phytocoenosis refers to floral community  Microbiocoenosis refers to microbial community.
  • 23. Community concept is one of the most important principles in Ecology.  This concept defines community as diverse organisms usually live together in an orderly manner.  Victor E. Shelford has defined “the community as an assemblage of populations living in a prescribed area or physical habitat with a definite trophic organization and metabolic pattern”. 
  • 24.    The individualistic hypothesis was proposed by H.A. Gleason( American Ecologist),1917. This hypothesis depicted a community as a chance assemblage of species found in an area because they have similar abiotic requirements. In other words ―vegetation is continuously variable in response to a continuously varying environment‖.
  • 25. The interactive hypothesis was proposed by F.E. Clements in 1916.  According to him, each community is an assemblage of closely linked species having mandatory biotic interactions that cause the community to function as an integrated unit. 
  • 26. Communities are stable, integrated, and orderly entities.  a. Biologically they have a highlypredictable composition.  b. The diversity and abundance of a species in a particular community will be the same before and after a disturbance.  c. Communities develop by passing through a series of predictable stages, culminating in a stable climax community. 
  • 27. Communities are neither stable nor predictable.  a. Plant and animal communities are ephemeral associations of species that just happen to share similar climatic requirements.  b. It is largely a matter of chance whether a similar community develops in the same area after a disturbance occurs. 
  • 28. The number of species  The relative abundance of those species  The types of species  The number, types and strength of interactions among species (i.e., the structure of food webs) 
  • 29.  1.Tropical        rain forests They found at equator. Temperatures and rainfall are high and variation is low. Plants grow all year long. High productivity, and high amounts of aboveground biomass. Highest amounts of BIODIVERSITY on Earth. Extraordinary structural diversity. A multilayered tree canopy is intermingled with vines, epiphytes, shrubs and herbs.
  • 30.  2.Temperate      Forests Precipitation is high and relatively constant throughout the year. This abundance of moisture allows trees to dominate the landscape. Plants experience a seasonal period of dormancy. Productivity higher than deserts or grasslands, lower than tropical forests. Biodiversity is moderate.
  • 31.  3. Grasslands These areas are also called prairies or steppes.  Precipitation conditions are quite dry.  Temperatures are moderate but highly SEASONAL.  Plant growth occurs in wet and warm months  Grasslands can develop in forested regions if recurring fires burn out  Productivity is lower than in forests.  Grassland soil is highly fertile. 
  • 32.  4.Deserts Low annual precipitation  Low productivity  Individual plants are widely spaced due to the intense competition for water.  Desert species must cope with extreme temperatures and aridity. 
  • 33.   Pond community – there may be populations of aquatic plants, herbivorous organisms, carnivores and bacteria at the bottom. They are interdependent for food supplies and form a specific group.
  • 34.       Species diversity is a key feature of biological communities. Species diversity affects the stability and productivity of communities. A latitudinal gradient of species diversity exists for many taxa. Species diversity declines as latitude increases. High diversity in the tropics leads to high productivity. Diversity is positively correlated with stability.
  • 35. Each community type experiences a characteristic type, frequency, and severity of disturbance. This is known as a community’s disturbance regime.  Disturbance patterns profoundly affect the composition of communities.  Examples—Fires in boreal forests and tree falls in temperate and tropical forests. 
  • 36. Definition •Community succession is defined as a gradual, continuous and unidirectional change in the species composition of a natural community over a long time.
  • 37. All ecosystems change over time  There are two types of ecosystem changes   Nondirectional directional changes-ReplacementFluctuations  Directional changes-Biogeographic change- Succession  All these types of changes may occur simultaneously within a single ecosystem
  • 39. Early stages Middle stages Final stages • Low biomass • Low nutrient levels • Low Species diversity • Short food chain • High productivity • Rich biomass • Rich organic nutrients • More species diversity • Stable high biomass • High organic nutrients & low productivity • Complex food chain • High species diversity &competition
  • 42. Increase in Species composition And species diversity Increase in species density And heterogeneity Increase in Organic nutrients Metabolic stability in Community metabolism
  • 43.
  • 44. Foundation species are considered the ―base‖ of a community, having the greatest influence on its overall structure.  They are usually the primary producers:  Foundation species may physically modify the environment to produce and maintain habitats that benefit the other organisms that use them.  E.g. the photosynthetic corals of the coral reef. 
  • 45.      A species whose presence is key to maintaining biodiversity within an ecosystem and to upholding an ecological community’s structure. The species may be a strong interactor or a weak interactor. Strong interactors are fewer in number and critical to the ecosystem function. Elimination of a keystone species may result in the loss of many other species in a community. E.g. beavers of Canada, Bengal tiger of India, African elephants.
  • 46. Invasive species are non-indigenous species whose introduction can cause harm to the community and the environment.  Invasive species are often better competitors than native species, resulting in population explosions.  These new species usually overtake the native populations, driving them to localized extinctions. 
  • 47.
  • 48. A niche is the role of an organism in a community (Charles Elton 1927)- functional niche = functional status of an organism in its community.  Joseph Grinnell (1917)referred niche as an organism’s physical environment- spatial niche  Niche is a property of a biotic community.  Niches includes all the physical, chemical and biological conditions.  The niche concept was formulated to study the role of organisms in community. 
  • 49. An edge is where two or more different vegetational communities meet. e.g. forestgrass land.  An ecotone is where two or more communities not only meet but also intergrade.  Edges result from abrupt changes in soil type, topographic differences, geomorphic differences and microclimatic changes.  Ecotones arise from the blending of two or more vegetational types. 
  • 50. A community is a complex group of individuals interacting and sharing an environment.  Communities can be characterized by their structure (the types and numbers of species present) and dynamics (how communities change over time). 
  • 51.      Dr. B. Victor is a highly experienced postgraduate professor, retired from the reputed educational institution - St. Xavier’ s College(Autonomous), Palayamkottai, India-627001. He was the dean of sciences, assistant controller of examinations and coordinator several academic research workshops. He has more than 32 years of teaching and research experience He has taught a diversity of courses and published 45 research articles in reputed national and international journals. Send your comments to : bonfiliusvictor@gmail.com