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ASSIGNMENT ON PBG- 513 
TOPIC – Plant Breeding , Its Objectives And Historical 
Development- Pre And Post Mendelian Era 
Presented by:- 
Avinash Gupta 
M.Sc. (pre.) 
Plant breeding & genetics 
RCA , MPUAT, Udaipur
CONTENTS 
1. Definition and Nature 
2. Role of Plant Breeding 
3. Challenges before plant breeder 
4. Objectives of plant breeding 
5. Activities in Plant Breeding 
6. History of Plant Breeding
Plant Breeding:- 
Plant breeding is the art, science and technology of changing the heredity of 
plants for human welfare. 
Nature of Plant Breeding:- 
1. Art 
•In earlier days man depends on his skills and judgement in 
selecting better plants. He knew nothing about the inheritance of 
characters, role of environment in producing them and the basis of 
variation in various plant characters. His method of selection was 
designed without the understanding of principles of inheritance. 
•Therefore during primitive time plant breeding was largely an art 
and very less science was involved in that. Even today success of 
selection depends upon ability of the person involved in the 
selection.
2. Science 
•Plant breeding is considered as the current phase of crop evolution. As the 
knowledge of genetics and other related science progresses plant breeding become 
less art and more science. 
•Especially, discovery of Mendel̕s 
work in 1900 added a lot to the knowledge of 
science. 
•Selection of desirable plant even today is an art it depends on the skill of a 
person. But alone skill is not enough, modern plant breeding is a combined effort 
of art and understanding and use of genetic principles. 
3. Technology 
Product of all plant breeding activities, whether dependent on the art or science, is 
improved variety, hybrids, synthetics and composites. This product is utilized by 
farmers for commercial cultivation. 
Therefore, plant breeding can be rightly viewed as a technology since it 
generates a useful product.
Role of Plant Breeding:- 
Human beings are dependent on the plants for: 
1. Food :- Breeding of field crops provides us food either directly (food 
grains) or indirectly (meat and milk). 
2. Shelter :- In addition to food by produce of agriculture farms are used in 
making shelter by farmers of rural areas. 
3. Clothing :- Breeding for fibre crops like cotton provides clothes for the 
human population. 
4. Fuels :- Crops like Euphorbia and Jatropha are used for Biofuel 
production. Breeding of such crops tackles the problems of energy 
production for rapidly increasing human population. Now a days , Maize is 
also used as an important source of Ethanol production.
5. Drugs :- Breeding of medicinal plants plays an important role in 
production of many important drugs. These drugs are used for treatment 
of various human and animal diseases. 
6. Entertainment:- Flowers play an essential role in peoples celebrations 
and everyday lives like weddings, Christmas etc. most of the medicinal 
plants are seasonal in nature. Shifting the seasonal timing of reproduction 
is a major goal of plant breeding efforts to produce novel varieties that 
are better adapted to local environments and changing climatic conditions.
Challenges before Plant Breeder :- 
1. Increasing population:- At present, the world population stand at 6.3 
billion and will reach at 10-12 billion during the next 50-70 years. The main 
problem from breeding respect is that the population is growing faster than 
increases in food productivity, to reduce the use of harmful agrochemicals 
and to produce nutritious and healthful food is greater today. 
2. Squeezing arable land :- Day-by-day the total arable land for 
agriculture is decreasing due to urbanization and industrial development. 
Breeders have to tackle this problem by releasing improved varieties of 
major crops which gives better production per unit area.
3. Erratic rainfall :- In India, rainfall is erratic, unpredictable and unevenly 
distributed. Over 80% of the annual rainfall is received in the four rainy 
months of June to September. Therefore, varieties which can tolerate dry 
spells and perform better at low water availability are needed to be develop 
by Indian Breeders. 
4. Mechanization:- The variety developed by plant breeders should give 
response to application of fertilizers, manures, irrigation and should be 
suitable for mechanical cultivation .
Objectives of Plant Breeding:- 
The prime objective of plant breeding is to develop superior plants over the 
existing ones in relation to their economic use. The objectives of plant breeding 
differ from crop to crop. A brief account of some important objectives are- 
1. Higher productivity- Increased yield has been the ultimate aim of most 
plant breeders. This can be achieved by developing more efficient genotypes 
having greater physiological efficiency. 
2. Improved quality- Improved quality of agricultural products has 
contributed a lot to the human well-being. Quality characters vary from one 
crop to another crop. For example, Grain size, colour , milling, and baking 
qualities in wheat (Triticum aestivum). 
3. Disease and Insect Resistance- Resistance varieties offer the 
cheapest and most convenient method of disease and insect management. In 
some cases, they offer only feasible means of control. eg. Rust inWheat.
4. Varieties for new seasons- The varieties for new seasons have been 
developed by adjusting the growth cycle of the variety to suit better to the 
available growing season. Traditionally maize is a kharif crop but, scientists 
are now able to grow maize throughout the year. Similarly, Mung is now 
grown as a summer crop in addition to the main kharif crop. 
5. Modification of agronomic characteristics- modification of 
agronomic characteristics such as plant height, tillering, branching, erect or 
trailing habit etc. is often desirable. For example, dwarfness in cereals is 
generally associated with lodging resistance and fertilizer responsiveness. 
6. Change in maturity duration- It permits new crop rotations and 
often extends the crop area. Development of wheat varieties suitable for late 
planting has permitted rice-wheat rotation. This objective is more desirable 
especially in those areas where multiple cropping system has been followed.
7. Photo and thermo insensitivity- Development of photo and 
thermo insensitive wheat and photo insensitive rice varieties has permitted 
their cultivation in new areas. eg. Cultivation of wheat in Kerala and W.B., 
cultivation of rice in Punjab and Himachal Pradesh.. 
8. Synchronous maturity- Synchronous maturity is highly desirable 
in crops where several pickings are necessary. Eg. Mungbean, pigeon pea, 
cotton etc. 
9. Non-shattering characteristics- It would be of great value in 
crops like mung, castor, soybean etc. where shattering is a major problem 
in case of many commercial varieties. 
10. Determinate growth- Development of varieties with determinate 
growth is desirable in crops like mung, pigeon pea, cotton, etc.
11. Dormancy- Dormancy plays both beneficial and harmful role according 
to the need of grower. For example, if we want next crop immediate after 
harvesting of previous crop, in such case dormancy is not required. But if we 
want to store the seed for its future purpose, a period of dormancy is essential. 
12. Elimination of toxic substances:- some crops have toxic substances 
which must be eliminated to make them safe for consumption. For example, 
• Khesari (Lathyrus odoratus) seeds have a neurotoxin, β- N- oxalyl - α-β- 
diaminopropionic acid (BOAA) that causes paralysis in humans. 
•Similarly, elimination of Erusic acid from Brassica oil and Gossypol from seed 
cotton is necessary to make them fit for consumption.
Activities in plant breeding: 
1.Creation of variation :-Variation means differences among 
individuals of a population or species for a specific character. Genetic 
variation is the source of raw material for selection. These are heritable 
and are transmitted from one generation to other. Such variation is useful 
in selection. Success of a breeding programme usually depends on the 
desired genetic variation. It can be done in following ways i.e. 
domestication, germplasm collection, plant introduction, hybridization, 
polyploidy, mutation, somaclonal variation and genetic engineering.
2. Selection:- 
During selection, the individual plant or group of plants having the 
desired characters are picked up from a population eliminating the undesirable 
ones. Those plants are selected which are looking promising for the character 
on thee basis of phenotype. The selected plants are then allowed to grow for 
setting their seeds. Seeds are selected and again a new crop is developed. This 
process is repeated again and again till the desired result is achieved. Selection 
acts on the genetic variation present in a population and produces a new 
population with improved characters.
3. Evaluation 
The newly selected lines/strains/populations are tested for yield and other traits 
and their performance is compared with existing best varieties called Checks. If 
the new lines/strain/population shows superior performance to the checks, it is 
released and notified as a new variety. 
4. Multiplication 
This step concerns with large scale certified seed production of the released and 
notified variety. 
5. Distribution 
Certified seed is ultimately sold to the farmers who use it for commercial crop 
cultivation.
Creation 
Of 
variation 
selection 
Evaluation 
multiplication 
distribution 
Domestication 
Naturally existing 
variability 
Somaclonal variation 
Creation of 
new 
variability 
Activities in plant breeding 
Germplasm collection 
Plant Introduction 
Hybridization 
Mutation 
Polyploidy 
Genetic engineering
History of Plant Breeding:- 
In broad sense history of plant breeding can be divided into 4 parts 
1. Pre Mendelian era:- before 1900 
2. Mendelian era:- 1900 to 1920. 
3. Post Mendelian era:- 1921 to 1950 
4. Modern era:- after 1950
1. Pre Mendelian era:- 
•9000 BC First evidence of plant domestication in hills above Tigris River 
•5000 BC Agricultural communities exist in Mesopotamia 
•4000 BC Egyptians used yeast in wine and bread making 
•3000 BC Domestication complete for all important food crops in the old world 
•1000 BC Domestication complete for all important food crops in the new 
world. 
•700 BC Assyrians and Babylonians - Hand pollination of date palm 
•1665 Hooke (England) – 1st Described the cell and known as father of cell-biology. 
•1676 Millington - Anthers function as male organs 
•1694 Camerarius (Germany) - First to demonstrate sex in plants.
•1717 Fairchild – produced the first artificial hybrid, popularly known as 
Fairchild҆s mule, by crossing Carnation with SweetWilliam variety of Dianthus. 
•1753 Linnaeus - Published "Species Plantarum". Binomial nomenclature of 
plant taxonomy officially begins with his general list of plant species 
•1761-66 Koelreuter (Germany) - Demonstrated that hybrid offspring received 
traits from both parents and were intermediate in most traits. 
•1779 Knight - Emphasized the practical aspects of hybrids. 
•1801 Lamarck - given Theory of evolution through inheritance of acquired 
characters . 
•1819 Shirreff - Utilized pure line selection to develop a new oat cultivar 
(released in 1824), and a new wheat cultivar (released in 1832)
•1831 Brown - Discovered the eukaryotic cell nucleus 
•1837-38 Schleiden and Schwann - Developed the cell theory 
•1859-89 Darwin - Published "Origin of Species"; and noted inbreeding, 
sterility, and differences in reciprocal crosses 
•1866 Mendel - Published "Experiments in plant hybridization"; discovered unit 
factors (genes), segregation of F2, recombination of 2 or more genes, and 
dominance of one allele over another; formulated the laws of inheritance 
•1884 Strasburger - Demonstrated fertilization and showed the fusion of the 2 
nuclei to form the zygote 
•1899 Novaschin and Guignard - Discovered double fertilization of egg and 
endosperm 
•1899 Hopkins - Described ear to row selection method.
2. Mendelian era:- 
•1900 Correns (Germany), DeVries (Holland) and Von Tschermak 
(Austria) - Independently rediscovered Mendel҆s laws of heredity. 
•Bateson- Introduced the terms "allelomorph", "homozygote", "heterozygote 
", "F1" and "F2" 
•1902 DeVries (Holland) - Proposed the mutation theory of evolution on his 
experiments on Oenothera lamarckiana. 
•Biffen (England) – on his studies on inheritance of studies on disease 
resistance; found that stripe rust resistance was due to a single gene 
•1903 Johannsen - Developed the pure line theory of selection
•1904 Hannig - Contributed to the idea of embryo culture 
•1906 Bateson - Introduced the term "genetics“ 
•YULE – gave the initial idea of multiple factor hypothesis. 
•1908 Nilsson- Ehle – given the explanation of multiple factor hypothesis for 
Grain colour of Wheat. 
•1908-09 Hardy (England) and Weinberg (Germany) – independently given 
algebraic equation that describes the algebraic equation within a population, 
also known as Hardy-Weinberg law. 
•1914 Shull - Introduced the term "heterosis " 
•Blakeslee - Discovered trisomics in Datura.
3. Post Mendelian era:- 
•1927 Muller - Reported artificial mutations in animals by X-rays 
•1928 Stadler - Described the mutagenic effects of X-rays in barley 
•1929 McClintock - First to report and number 10 chromosomes in maize 
•1931 Stern, Creighton and McClintock - Provided the cytological proof of 
crossing-over 
•1933 Rhoades - Discovered cytoplasmic male sterility in maize 
•1944 Avery, MacLeod and McCarty - Described the transforming principle 
and suggested that DNA, not protein, is the hereditary material 
•1945 Hull - Proposed recurrent selection method 
•1946 Comstock et al. - Suggested reciprocal (half-sib) recurrent selection 
•1950 McClintock - Described the Ac-Ds system of transposable elements in 
maize.
4. Modern era:- 
•1952 Jensen - 1st suggested the use of multilines in Oats. 
•1953 N.E.Borlaug – 1st outlined the method of developing multilines in 
wheat. 
•1955 Benzer – Based on his work on rII locus of T4 Bacteriophage he 
gave the subdivisions of genes in cistron, recon and muton. 
•1963 Vanderplank – developed the concept of vertical and horizontal 
resistance. 
•1964 N.E.Borlaug – developed the high yielding dwarf varieties of Wheat 
which resulted in green revolution.
•1968 Donald – developed the concept of crop ideotype in wheat. 
•1978 - Development of worlds 1st Rice hybrid (CMS based) for commercial 
cultivation in China. 
•1983 – Development of 1st transgenic (genetically engineered) plant of Tobacco 
in U.S.A. 
•1987 – Development of 1st transgenic cotton plant by Monsanto company in 
U.S.A. 
•1994 – Flavr Savr tomato was introduced by Calgene company of California. It 
was the 1st commercially grown genetically engineered crop.
History of Plant Breeding in India 
•1871- Government of India created the Department of Agriculture. 
•1905- The Imperial Agricultural Research Institute was established in Pusa 
(Bihar), this was the first Agricultural Research Institute in the country. 
•1921- The Indian Central cotton committee was established. 
•1929- Imperial Council of Agricultural Research was established in New Delhi. 
•1936- Imperial Agricultural Research Institute shifted to its present location in 
New Delhi. 
•1956- Project for Intensification of Regional Research on Cotton, Oilseeds and 
Millets (PIRCOM) was initiated in order to intensify research on these crops.
•1957- All India Coordinated Maize Improvement Project was started with 
the objective of exploiting Heterosis. 
•1961- First Maize hybrid was released (Ganga1, Ganga101, Deccan and 
Ranjit) 
•1964- First Sorghum hybrid (CSH-1) was released. 
•1965- First Bajra hybrid (HB-1) was released. 
•1991 – 1st pigeon pea hybrid (ICPH-8) was released from ICRISAT, 
Hyderabad
Some indian plant breeders:- 
•T. S. Venkatraman - he transferred thick stem and high sugar contents from 
tropical cane ( Saccharum officinarum) to indian cane (Saccharum barberi). 
this process is known as Noblisation of sugarcane. 
•B. P. PAL – 1st director general of ICAR. He developed some superior 
disease resistance variety of Wheat. 
•M. S. Swaminathan – father of green revolution in India. He developed the 
high yielding dwarf varieties of Wheat. 
•Puskarnath – a famous potato breeder who developed several high yielding 
varieties of potato. 
•N. G. P. Rao – he is an eminent sorghum breeder and developed the worlds 
1st hybrid of sorghum (CSH-1).
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Plant breeding, its objective and historical development- pre and post mendelian era01 avinash

  • 1. ASSIGNMENT ON PBG- 513 TOPIC – Plant Breeding , Its Objectives And Historical Development- Pre And Post Mendelian Era Presented by:- Avinash Gupta M.Sc. (pre.) Plant breeding & genetics RCA , MPUAT, Udaipur
  • 2. CONTENTS 1. Definition and Nature 2. Role of Plant Breeding 3. Challenges before plant breeder 4. Objectives of plant breeding 5. Activities in Plant Breeding 6. History of Plant Breeding
  • 3. Plant Breeding:- Plant breeding is the art, science and technology of changing the heredity of plants for human welfare. Nature of Plant Breeding:- 1. Art •In earlier days man depends on his skills and judgement in selecting better plants. He knew nothing about the inheritance of characters, role of environment in producing them and the basis of variation in various plant characters. His method of selection was designed without the understanding of principles of inheritance. •Therefore during primitive time plant breeding was largely an art and very less science was involved in that. Even today success of selection depends upon ability of the person involved in the selection.
  • 4. 2. Science •Plant breeding is considered as the current phase of crop evolution. As the knowledge of genetics and other related science progresses plant breeding become less art and more science. •Especially, discovery of Mendel̕s work in 1900 added a lot to the knowledge of science. •Selection of desirable plant even today is an art it depends on the skill of a person. But alone skill is not enough, modern plant breeding is a combined effort of art and understanding and use of genetic principles. 3. Technology Product of all plant breeding activities, whether dependent on the art or science, is improved variety, hybrids, synthetics and composites. This product is utilized by farmers for commercial cultivation. Therefore, plant breeding can be rightly viewed as a technology since it generates a useful product.
  • 5. Role of Plant Breeding:- Human beings are dependent on the plants for: 1. Food :- Breeding of field crops provides us food either directly (food grains) or indirectly (meat and milk). 2. Shelter :- In addition to food by produce of agriculture farms are used in making shelter by farmers of rural areas. 3. Clothing :- Breeding for fibre crops like cotton provides clothes for the human population. 4. Fuels :- Crops like Euphorbia and Jatropha are used for Biofuel production. Breeding of such crops tackles the problems of energy production for rapidly increasing human population. Now a days , Maize is also used as an important source of Ethanol production.
  • 6. 5. Drugs :- Breeding of medicinal plants plays an important role in production of many important drugs. These drugs are used for treatment of various human and animal diseases. 6. Entertainment:- Flowers play an essential role in peoples celebrations and everyday lives like weddings, Christmas etc. most of the medicinal plants are seasonal in nature. Shifting the seasonal timing of reproduction is a major goal of plant breeding efforts to produce novel varieties that are better adapted to local environments and changing climatic conditions.
  • 7. Challenges before Plant Breeder :- 1. Increasing population:- At present, the world population stand at 6.3 billion and will reach at 10-12 billion during the next 50-70 years. The main problem from breeding respect is that the population is growing faster than increases in food productivity, to reduce the use of harmful agrochemicals and to produce nutritious and healthful food is greater today. 2. Squeezing arable land :- Day-by-day the total arable land for agriculture is decreasing due to urbanization and industrial development. Breeders have to tackle this problem by releasing improved varieties of major crops which gives better production per unit area.
  • 8. 3. Erratic rainfall :- In India, rainfall is erratic, unpredictable and unevenly distributed. Over 80% of the annual rainfall is received in the four rainy months of June to September. Therefore, varieties which can tolerate dry spells and perform better at low water availability are needed to be develop by Indian Breeders. 4. Mechanization:- The variety developed by plant breeders should give response to application of fertilizers, manures, irrigation and should be suitable for mechanical cultivation .
  • 9. Objectives of Plant Breeding:- The prime objective of plant breeding is to develop superior plants over the existing ones in relation to their economic use. The objectives of plant breeding differ from crop to crop. A brief account of some important objectives are- 1. Higher productivity- Increased yield has been the ultimate aim of most plant breeders. This can be achieved by developing more efficient genotypes having greater physiological efficiency. 2. Improved quality- Improved quality of agricultural products has contributed a lot to the human well-being. Quality characters vary from one crop to another crop. For example, Grain size, colour , milling, and baking qualities in wheat (Triticum aestivum). 3. Disease and Insect Resistance- Resistance varieties offer the cheapest and most convenient method of disease and insect management. In some cases, they offer only feasible means of control. eg. Rust inWheat.
  • 10. 4. Varieties for new seasons- The varieties for new seasons have been developed by adjusting the growth cycle of the variety to suit better to the available growing season. Traditionally maize is a kharif crop but, scientists are now able to grow maize throughout the year. Similarly, Mung is now grown as a summer crop in addition to the main kharif crop. 5. Modification of agronomic characteristics- modification of agronomic characteristics such as plant height, tillering, branching, erect or trailing habit etc. is often desirable. For example, dwarfness in cereals is generally associated with lodging resistance and fertilizer responsiveness. 6. Change in maturity duration- It permits new crop rotations and often extends the crop area. Development of wheat varieties suitable for late planting has permitted rice-wheat rotation. This objective is more desirable especially in those areas where multiple cropping system has been followed.
  • 11. 7. Photo and thermo insensitivity- Development of photo and thermo insensitive wheat and photo insensitive rice varieties has permitted their cultivation in new areas. eg. Cultivation of wheat in Kerala and W.B., cultivation of rice in Punjab and Himachal Pradesh.. 8. Synchronous maturity- Synchronous maturity is highly desirable in crops where several pickings are necessary. Eg. Mungbean, pigeon pea, cotton etc. 9. Non-shattering characteristics- It would be of great value in crops like mung, castor, soybean etc. where shattering is a major problem in case of many commercial varieties. 10. Determinate growth- Development of varieties with determinate growth is desirable in crops like mung, pigeon pea, cotton, etc.
  • 12. 11. Dormancy- Dormancy plays both beneficial and harmful role according to the need of grower. For example, if we want next crop immediate after harvesting of previous crop, in such case dormancy is not required. But if we want to store the seed for its future purpose, a period of dormancy is essential. 12. Elimination of toxic substances:- some crops have toxic substances which must be eliminated to make them safe for consumption. For example, • Khesari (Lathyrus odoratus) seeds have a neurotoxin, β- N- oxalyl - α-β- diaminopropionic acid (BOAA) that causes paralysis in humans. •Similarly, elimination of Erusic acid from Brassica oil and Gossypol from seed cotton is necessary to make them fit for consumption.
  • 13. Activities in plant breeding: 1.Creation of variation :-Variation means differences among individuals of a population or species for a specific character. Genetic variation is the source of raw material for selection. These are heritable and are transmitted from one generation to other. Such variation is useful in selection. Success of a breeding programme usually depends on the desired genetic variation. It can be done in following ways i.e. domestication, germplasm collection, plant introduction, hybridization, polyploidy, mutation, somaclonal variation and genetic engineering.
  • 14. 2. Selection:- During selection, the individual plant or group of plants having the desired characters are picked up from a population eliminating the undesirable ones. Those plants are selected which are looking promising for the character on thee basis of phenotype. The selected plants are then allowed to grow for setting their seeds. Seeds are selected and again a new crop is developed. This process is repeated again and again till the desired result is achieved. Selection acts on the genetic variation present in a population and produces a new population with improved characters.
  • 15. 3. Evaluation The newly selected lines/strains/populations are tested for yield and other traits and their performance is compared with existing best varieties called Checks. If the new lines/strain/population shows superior performance to the checks, it is released and notified as a new variety. 4. Multiplication This step concerns with large scale certified seed production of the released and notified variety. 5. Distribution Certified seed is ultimately sold to the farmers who use it for commercial crop cultivation.
  • 16. Creation Of variation selection Evaluation multiplication distribution Domestication Naturally existing variability Somaclonal variation Creation of new variability Activities in plant breeding Germplasm collection Plant Introduction Hybridization Mutation Polyploidy Genetic engineering
  • 17. History of Plant Breeding:- In broad sense history of plant breeding can be divided into 4 parts 1. Pre Mendelian era:- before 1900 2. Mendelian era:- 1900 to 1920. 3. Post Mendelian era:- 1921 to 1950 4. Modern era:- after 1950
  • 18. 1. Pre Mendelian era:- •9000 BC First evidence of plant domestication in hills above Tigris River •5000 BC Agricultural communities exist in Mesopotamia •4000 BC Egyptians used yeast in wine and bread making •3000 BC Domestication complete for all important food crops in the old world •1000 BC Domestication complete for all important food crops in the new world. •700 BC Assyrians and Babylonians - Hand pollination of date palm •1665 Hooke (England) – 1st Described the cell and known as father of cell-biology. •1676 Millington - Anthers function as male organs •1694 Camerarius (Germany) - First to demonstrate sex in plants.
  • 19. •1717 Fairchild – produced the first artificial hybrid, popularly known as Fairchild҆s mule, by crossing Carnation with SweetWilliam variety of Dianthus. •1753 Linnaeus - Published "Species Plantarum". Binomial nomenclature of plant taxonomy officially begins with his general list of plant species •1761-66 Koelreuter (Germany) - Demonstrated that hybrid offspring received traits from both parents and were intermediate in most traits. •1779 Knight - Emphasized the practical aspects of hybrids. •1801 Lamarck - given Theory of evolution through inheritance of acquired characters . •1819 Shirreff - Utilized pure line selection to develop a new oat cultivar (released in 1824), and a new wheat cultivar (released in 1832)
  • 20. •1831 Brown - Discovered the eukaryotic cell nucleus •1837-38 Schleiden and Schwann - Developed the cell theory •1859-89 Darwin - Published "Origin of Species"; and noted inbreeding, sterility, and differences in reciprocal crosses •1866 Mendel - Published "Experiments in plant hybridization"; discovered unit factors (genes), segregation of F2, recombination of 2 or more genes, and dominance of one allele over another; formulated the laws of inheritance •1884 Strasburger - Demonstrated fertilization and showed the fusion of the 2 nuclei to form the zygote •1899 Novaschin and Guignard - Discovered double fertilization of egg and endosperm •1899 Hopkins - Described ear to row selection method.
  • 21. 2. Mendelian era:- •1900 Correns (Germany), DeVries (Holland) and Von Tschermak (Austria) - Independently rediscovered Mendel҆s laws of heredity. •Bateson- Introduced the terms "allelomorph", "homozygote", "heterozygote ", "F1" and "F2" •1902 DeVries (Holland) - Proposed the mutation theory of evolution on his experiments on Oenothera lamarckiana. •Biffen (England) – on his studies on inheritance of studies on disease resistance; found that stripe rust resistance was due to a single gene •1903 Johannsen - Developed the pure line theory of selection
  • 22. •1904 Hannig - Contributed to the idea of embryo culture •1906 Bateson - Introduced the term "genetics“ •YULE – gave the initial idea of multiple factor hypothesis. •1908 Nilsson- Ehle – given the explanation of multiple factor hypothesis for Grain colour of Wheat. •1908-09 Hardy (England) and Weinberg (Germany) – independently given algebraic equation that describes the algebraic equation within a population, also known as Hardy-Weinberg law. •1914 Shull - Introduced the term "heterosis " •Blakeslee - Discovered trisomics in Datura.
  • 23. 3. Post Mendelian era:- •1927 Muller - Reported artificial mutations in animals by X-rays •1928 Stadler - Described the mutagenic effects of X-rays in barley •1929 McClintock - First to report and number 10 chromosomes in maize •1931 Stern, Creighton and McClintock - Provided the cytological proof of crossing-over •1933 Rhoades - Discovered cytoplasmic male sterility in maize •1944 Avery, MacLeod and McCarty - Described the transforming principle and suggested that DNA, not protein, is the hereditary material •1945 Hull - Proposed recurrent selection method •1946 Comstock et al. - Suggested reciprocal (half-sib) recurrent selection •1950 McClintock - Described the Ac-Ds system of transposable elements in maize.
  • 24. 4. Modern era:- •1952 Jensen - 1st suggested the use of multilines in Oats. •1953 N.E.Borlaug – 1st outlined the method of developing multilines in wheat. •1955 Benzer – Based on his work on rII locus of T4 Bacteriophage he gave the subdivisions of genes in cistron, recon and muton. •1963 Vanderplank – developed the concept of vertical and horizontal resistance. •1964 N.E.Borlaug – developed the high yielding dwarf varieties of Wheat which resulted in green revolution.
  • 25. •1968 Donald – developed the concept of crop ideotype in wheat. •1978 - Development of worlds 1st Rice hybrid (CMS based) for commercial cultivation in China. •1983 – Development of 1st transgenic (genetically engineered) plant of Tobacco in U.S.A. •1987 – Development of 1st transgenic cotton plant by Monsanto company in U.S.A. •1994 – Flavr Savr tomato was introduced by Calgene company of California. It was the 1st commercially grown genetically engineered crop.
  • 26. History of Plant Breeding in India •1871- Government of India created the Department of Agriculture. •1905- The Imperial Agricultural Research Institute was established in Pusa (Bihar), this was the first Agricultural Research Institute in the country. •1921- The Indian Central cotton committee was established. •1929- Imperial Council of Agricultural Research was established in New Delhi. •1936- Imperial Agricultural Research Institute shifted to its present location in New Delhi. •1956- Project for Intensification of Regional Research on Cotton, Oilseeds and Millets (PIRCOM) was initiated in order to intensify research on these crops.
  • 27. •1957- All India Coordinated Maize Improvement Project was started with the objective of exploiting Heterosis. •1961- First Maize hybrid was released (Ganga1, Ganga101, Deccan and Ranjit) •1964- First Sorghum hybrid (CSH-1) was released. •1965- First Bajra hybrid (HB-1) was released. •1991 – 1st pigeon pea hybrid (ICPH-8) was released from ICRISAT, Hyderabad
  • 28. Some indian plant breeders:- •T. S. Venkatraman - he transferred thick stem and high sugar contents from tropical cane ( Saccharum officinarum) to indian cane (Saccharum barberi). this process is known as Noblisation of sugarcane. •B. P. PAL – 1st director general of ICAR. He developed some superior disease resistance variety of Wheat. •M. S. Swaminathan – father of green revolution in India. He developed the high yielding dwarf varieties of Wheat. •Puskarnath – a famous potato breeder who developed several high yielding varieties of potato. •N. G. P. Rao – he is an eminent sorghum breeder and developed the worlds 1st hybrid of sorghum (CSH-1).